If you want to apply for scholarships please fill up the form below (both PhD students and Postdocs can apply).

Please be aware that these scholarships are in the form of tuition fee waivers only, they do not cover traveling and lodging.

You need to register for the course before applying for the scholarship (at the course webpage). If you are applying for a scholarship you will not be expected to pay the course, and if you do not get granted you can always withdraw.


  1. Funding for their PhD. Students  with no funding to develop their PhD will receive the maximum 4 points. Part-time research grants or contracts (teaching assistantships) receive 2 points. Students with full financial support to develop their PhD provided by their university or an external institution / agency will receive 0 points. The student’s PhD organization or academic advisor must certify the situation. 4 point scale.
  2. Belonging to developing countries1 point.
  3. A short project description (maximum 300 words) related to the course or workshop topic. 5 point scale.


August 10th-12th, 2020


In this workshop students will be introduced to the ENMTools R package, which is a package for constructing niche and distribution models and using them to test hypotheses about patterns of niche evolution.  ENMTools contains a streamlined workflow for making, visualizing, and projecting models, as well as new methods for evaluating model predictions, statistical significance, and biases in the model construction and projection process.

It also implements Monte Carlo tests addressing many fundamental questions regarding niche evolution, such as:

  • Are two species’ environmental associations identical?
  • Are two species’ environmental associations more or less similar than expected based on the available habitat in their ranges?
  • Are biogeographic boundaries determined by environmental gradients?
  • What are the patterns of niche evolution across clades?


Graduate or postgraduate degree in any Biosciences discipline. All participants must have a personal computer (Windows, Macintosh), with webcam if possible, headphones, and a good internet connection.

This is an advanced course intended for students who are already familiar with ENM/SDM theory and are used to working in R.



This course will be delivered online.

Please check the schedule for the live online part, and be aware that it is GMT+1.


August 10th-12th, 2020




15 hours online.


Places are limited to 14 participants and will be occupied by strict registration order.

Participants who have completed the course will receive a certificate at the end of it.

Dan Warren instructor for Transmitting Science

Dr. Dan Warren

Senckenberg Biodiversity and Climate Research Center

Haris Saslis coordinator for Transmitting Science

Dr. Haris Saslis

Transmitting Science


Before the course (online tutorials)

  • Installing packages
  • Spatial data in R
  • Point data, vector data, and raster data
  • GBIF, the Global Biodiversity Information Facility


  • Building ENMTools species objects
  • ENMs with ENMTools
  • Bioclim
  • Domain
  • GLM
  • GAM
  • Maxent
  • Random forest
  • Boosted regression trees
  • Poisson point process models
  • Hypervolume
  • Projecting models across space and time


  • Model selection overview
  • Discrimination metrics (geographic space)
  • Discrimination metrics (environment space)
  • Calibration metrics
  • Monte Carlo methods: conceptual overview
  • Raes and ter Steege and Bohl et al. methods for estimating ENM significance
  • Warren et al. (in review) method for measuring methodological bias


  • Measuring niche breadth on ENMs
  • Measuring niche overlap on ENMs
  • Identity tests
  • Background tests
  • Rangebreak tests
  • Ecospat tests via ENMTools
  • Pairwise MOSES tests for taxonomic scale
  • Age-range correlation
  • Age-overlap correlation


  • Course Fee
  • Early bird (until July 31st, 2020):
  • 240 € *(192 € for Ambassador Institutions)
  • Regular (after July 31st, 2020):
  • 285 € *(228 € for Ambassador Institutions)
  • This includes course material (VAT included).
    * Participants from companies/industry will have an extra charge of 100 €.

You can check the list of Ambassador Institutions. If you want your institution to become a Transmitting Science Ambassador please contact us at


  • Monday to Wednesday (GMT+1):
    • 9:00 to 12:00 online live lessons

The rest of the time will be taught with recorded classes, to watch before the course and between the live sessions.


Discounts are not cumulative and apply only on the Course Fee. We offer the possibility of paying in two instalments (contact the course coordinator).


Terms and Conditions and Privacy Policy: In order to process the registration, management and maintenance of your profile as course participant on our website, we need your personal data. If you want more information about the processing of your data and your rights, we recommend that you read our privacy policy carefully (below).

  • After registration you will receive confirmation of your acceptance in the course. Payment is not required during registration.

  • Preferred names will be used for identifiers and communication. Legal names will be used for certificates and invoices.
  • We will use your phone number only in case we need to reach you just before or during the course.
  • Course Fee (€)

  • Discounts

    (Discounts are not cumulative and apply only on the Course Fee, not on the Accommodation Package or other options).
  • Donate for grants

    We accept donations, which would be entirely devoted to support students without financial resources so that they may offset the cost of participating in our courses and workshops. More information about donations and our transparency policy here.
    In case you want to make a donation, you may add it to your Course Fee. In that case please select the option below:
  • The amount due is (€)

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    Basic information about Protection of User Data that contacts through the Web: Controller is TRANSMITTING SCIENCE SL, to attend and answer the communications or requests received. The legitimate interest of TRANSMITTING SCIENCE SL in attending your queries. Recipients are housing services and IT consulting services. International Data Transfers are not planned. Access, rectify and delete data, as well as other rights, as explained in the additional information. You can access detailed information on the processing on our privacy policy, legal notice and cookie policy in our website.
  • Cancellation policy:

    If you wish to cancel your participation in this course, cancellations up to 20 days before the course start date will incur a 30 % cancellation Course Fee. For later cancellations, or non-attendance, 75 % of the Course Fee will be charged.
    If Transmitting Science must cancel this course due to unforeseen circumstances beyond the control of Transmitting Science, you will either be entitled to a full refund of the Course Fee, or your Course Fee can be credited toward a future course. Transmitting Science is not responsible for travel fees, or any expenses incurred by you as a result of such cancellation. Every effort will be made to avoid the cancellation of any confirmed course.
Publicado en Convocatorias.

Cuba, La Habana. Investigador Titular del Centro de Investigaciones Pesqueras, doctor en Ciencias en el Uso, Manejo y Preservación de los Recursos, y maestro en Ciencias del Agua.

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